$re = '/^(\S+)[^\S\r\n]+/m';
$str = 'GAXI01000525.151.1950 Eukaryota;Opisthokonta;Holozoa;Metazoa (Animalia);Eumetazoa;Bilateria;Arthropoda;Hexapoda;Ellipura;Collembola;Tetrodontophora bielanensis (giant springtail)
GAXI01000526.151.1950 Eukaryota;Opisthokonta;Holozoa;Metazoa (Animalia);Eumetazoa;Bilateria;Arthropoda;Hexapoda;Ellipura;Collembola;Tetrodontophora bielanensis (giant springtail)
GAXI01005455.1.1233 Bacteria;Bacteroidetes;Flavobacteriia;Flavobacteriales;Flavobacteriaceae;Chryseobacterium;Tetrodontophora bielanensis (giant springtail)
GAXI01006199.29.1525 Bacteria;Chlamydiae;Chlamydiae;Chlamydiales;Simkaniaceae;Candidatus Rhabdochlamydia;Tetrodontophora bielanensis (giant springtail)';
$subst = "$1;";
$result = preg_replace($re, $subst, $str);
echo "The result of the substitution is ".$result;
Please keep in mind that these code samples are automatically generated and are not guaranteed to work. If you find any syntax errors, feel free to submit a bug report. For a full regex reference for PHP, please visit: http://php.net/manual/en/ref.pcre.php