^start of line
Negative Lookahead(?!(\d)(?:-?\1)*$) Non-Capturing Group(?:-?\1)* *repeats the group contents below any number of times:
-?optionally matches the literal - (4510 / 558 / 2D16)
\1the text saved by group 1
$end of line
\d{2}a digit, repeated exactly 2 times
-?optionally matches the literal - (4510 / 558 / 2D16)
\d{7}a digit, repeated exactly 7 times
$end of line
g modifier: global. Finds all matches instead of stopping after the first
m modifier: multiline. Causes ^ and $ to match the start and end of each line, not only the start and end of the string