use strict;
my $str = '<!DOCTYPE html>
<html>
<body>
<h1>My Test Heading</h1>
<a href>text1 text2</a>
<% testfunction() %>
<p>My Test paragraph.</p>
</body>
</html>';
my $regex = qr/(?<=[>])\s+(?=[<])/p;
my $subst = '';
my $result = $str =~ s/$regex/$subst/rg;
print "The result of the substitution is' $result\n";
Please keep in mind that these code samples are automatically generated and are not guaranteed to work. If you find any syntax errors, feel free to submit a bug report. For a full regex reference for Perl, please visit: http://perldoc.perl.org/perlre.html