use strict;
my $str = '#| label: my_chunk_label
my_function_name <- function(x)';
my $regex = qr/(?:^#\|\h+label:\h+|\G(?!^))[^\r\n_]*\K_/mp;
my $subst = '-';
my $result = $str =~ s/$regex/$subst/rg;
print "The result of the substitution is' $result\n";
Please keep in mind that these code samples are automatically generated and are not guaranteed to work. If you find any syntax errors, feel free to submit a bug report. For a full regex reference for Perl, please visit: http://perldoc.perl.org/perlre.html